/* ---------- Reset & base ---------- */
* { box-sizing: border-box; margin: 0; padding: 0; }

:root {
  --font: "EB Garamond", Georgia, "Times New Roman", serif;

  --ink: #2b2532;
  --ink-soft: #55505c;
  --paper: #faf8f5;
  --paper-alt: #f2f1ee;
  --line: #e3dce9;

  --purple: #6d4695;
  --purple-dark: #452c60;
  --orange: #e08128;
  --orange-dark: #b8611a;
  --periwinkle: #7b7fe0;

  --maxw: 1040px;
  --textw: 700px;
}

html { scroll-behavior: smooth; font-size: 20px; }

body {
  font-family: var(--font);
  color: var(--ink);
  background: var(--paper);
  line-height: 1.7;
  font-size: 22px;
}

h1, h2, h3 {
  font-family: var(--font);
  font-weight: 500;
  line-height: 1.25;
  color: var(--ink);
}

h1 { font-size: 2.5rem; margin-bottom: 0.75rem; }
h2 { font-size: 1.9rem; margin-bottom: 1.1rem; }
h3 { font-size: 1.3rem; margin-bottom: 0.4rem; }

p { margin-bottom: 1.1rem; color: var(--ink-soft); }

a { color: var(--purple); text-decoration: none; }
a:hover { color: var(--orange-dark); text-decoration: underline; }

ul { margin-left: 1.2rem; margin-bottom: 1rem; color: var(--ink-soft); }
li { margin-bottom: 0.6rem; }

strong, b { font-weight: 600; color: var(--ink); }

.wrap {
  max-width: var(--maxw);
  margin: 0 auto;
  padding: 0 2rem;
}

/* ---------- Nav ---------- */
header.site-header {
  background: var(--paper);
  border-bottom: 1px solid #d8d3cd;
  box-shadow: 0 6px 12px -10px rgba(43,37,50,0.18);
  position: sticky;
  top: 0;
  z-index: 10;
}

/* Home page: logo centered above nav */
.logo-row {
  display: flex;
  justify-content: center;
  padding-top: 0.5rem;
}
.logo-row img { height: 140px; width: auto; }

.header-tagline {
  text-align: center;
  font-size: 1.1rem;
  font-weight: 700;
  color: var(--ink-soft);
  margin-top: -0.4rem;
}

nav.wrap {
  display: flex;
  align-items: center;
  justify-content: center;
  flex-wrap: wrap;
  padding-top: 0.6rem;
  padding-bottom: 1rem;
  gap: 0.75rem;
}

/* Inner pages: logo left, nav right */
.header-inner {
  display: flex;
  align-items: center;
  justify-content: space-between;
  flex-wrap: wrap;
  gap: 1rem;
  padding: 0.45rem 2rem;
}
.header-left {
  display: flex;
  flex-direction: column;
  align-items: flex-start;
}
.header-left img { height: 68px; width: auto; }

.nav-links {
  display: flex;
  gap: 2.75rem;
  flex-wrap: wrap;
  list-style: none;
  justify-content: center;
  margin: 0;
}
.nav-links li { margin-bottom: 0; }
.header-inner .nav-links { justify-content: flex-end; gap: 2.25rem; }

@media (max-width: 720px) {
  .header-inner { flex-direction: column; align-items: center; text-align: center; padding: 0.2rem 0.3rem; gap: 0.15rem; }
  .header-left { align-items: center; }
  .header-inner .nav-links { justify-content: center; flex-wrap: nowrap; gap: 0.45rem; }
  .header-inner .nav-links a { font-size: 0.78rem; font-weight: 700; white-space: nowrap; }
  .header-left img { height: 62px; }
}
.nav-links a {
  color: var(--ink);
  font-size: 1.25rem;
}
.nav-links a:hover, .nav-links a.active {
  color: var(--orange-dark);
  text-decoration: none;
  border-bottom: 1px solid var(--orange-dark);
}

.nav-links li.has-dropdown { position: relative; }
.dropdown-menu {
  display: none;
  position: absolute;
  top: 100%;
  left: 0;
  min-width: 170px;
  background: var(--paper);
  border-radius: 4px;
  box-shadow: 0 10px 24px -6px rgba(43,37,50,0.18);
  padding: 0.4rem 0;
  z-index: 20;
  list-style: none;
  margin: 0;
}
.has-dropdown:hover .dropdown-menu,
.has-dropdown:focus-within .dropdown-menu {
  display: block;
}
.dropdown-menu li { margin-bottom: 0; }
.dropdown-menu a {
  display: block;
  color: var(--ink) !important;
  font-size: 1.05rem;
  padding: 0.55rem 1.25rem;
  border-bottom: none !important;
}
.dropdown-menu a:hover, .dropdown-menu a.active {
  color: var(--orange-dark) !important;
  background: var(--paper-alt);
}

/* Inner pages (everything except home): purple header, tighter spacing */
.inner-page header.site-header {
  background: var(--purple-dark);
  border-bottom: none;
}
.inner-page .nav-links a { color: var(--paper); }
.inner-page .nav-links a:hover, .inner-page .nav-links a.active {
  color: var(--orange);
  border-bottom: 1px solid var(--orange);
}
.inner-page main { padding: 0.5rem 0 4.5rem; }
.inner-page section { margin-bottom: 4rem; }
.inner-page .hero.plain { padding: 1.75rem 0 0.25rem; margin-bottom: 0.5rem; }

/* Contact, Publications, Lab Values: much less vertical padding — these pages run short */
.page-tight main { padding: 0.5rem 0 1.5rem; }
.page-tight section { margin-bottom: 2rem; }

/* Bare hero: just an eyebrow label, no heading/lede — collapse it down to the label itself */
.inner-page .hero.plain.hero-bare { padding: 1rem 0 0; margin-bottom: 0.25rem; }
.hero.plain.hero-bare .eyebrow { margin-bottom: 0; }

/* ---------- Hero ---------- */
.hero {
  padding: 3rem 0 6.5rem;
  margin-bottom: 0;
  background: var(--paper-alt);
  text-align: center;
}
.hero.plain {
  background: transparent;
  padding: 4rem 0 1.5rem;
}
.hero .eyebrow {
  font-size: 1.2rem;
  font-style: normal;
  letter-spacing: 0.03em;
  color: var(--orange-dark);
  margin-bottom: 1.75rem;
}
.hero h1 {
  max-width: 880px;
  margin-left: auto;
  margin-right: auto;
  font-size: 2.1rem;
}
.hero .lede {
  font-size: 1.3rem;
  max-width: 780px;
  margin: 1.5rem auto 0;
  color: var(--ink);
}

.banner {
  background: var(--purple-dark);
  color: var(--paper);
}
.banner .wrap {
  padding: 0.85rem 2rem;
  display: flex;
  align-items: center;
  justify-content: space-between;
  flex-wrap: wrap;
  gap: 0.5rem;
  font-size: 0.95rem;
}
.banner a { color: var(--orange); font-weight: 600; text-decoration: underline; }

.band {
  background: var(--periwinkle);
  color: white;
  padding: 2.25rem 0;
  text-align: center;
}
.band h2, .band h3 { color: white; }
.band p { color: rgba(255,255,255,0.92); max-width: var(--textw); margin-left: auto; margin-right: auto; }
.band a { color: white; text-decoration: underline; }

/* ---------- Sections / cards ---------- */
main { padding: 1.5rem 0 6rem; }
section { margin-bottom: 6rem; }
section:last-child { margin-bottom: 0; }

.prose { max-width: var(--textw); margin: 0 auto; text-align: center; }
.prose p { max-width: none; }
.prose .direction-question {
  font-style: italic;
  font-size: 1.05rem;
  color: var(--orange-dark);
  margin-top: -0.5rem;
}

/* Research page: microscopy figures, sized consistently and centered within the prose */
.research-figure {
  margin: 2rem auto;
  max-width: none;
}
.research-figure img {
  width: 400px;
  height: 400px;
  object-fit: cover;
  border-radius: 4px;
  display: block;
  margin: 0 auto;
}
.research-figure figcaption {
  margin-top: 0.75rem;
  font-size: 0.85rem;
  font-style: italic;
  color: var(--ink-soft);
}

.btn {
  display: inline-block;
  padding: 0.7rem 1.7rem;
  background: var(--purple);
  color: var(--paper) !important;
  border-radius: 3px;
  font-size: 1rem;
  text-decoration: none !important;
}
.btn:hover { background: var(--purple-dark); }
.btn.secondary {
  background: transparent;
  color: var(--ink) !important;
  border: 1px solid var(--ink);
}
.btn.secondary:hover { background: var(--ink); color: var(--paper) !important; }
.cta-row { display: flex; gap: 1rem; flex-wrap: wrap; margin-top: 2rem; justify-content: center; }

.card {
  background: transparent;
  border-top: 1px solid var(--line);
  border-radius: 0;
  padding: 2.25rem 0 0;
}
.card:first-child { border-top: none; padding-top: 0; }

/* ---------- Question / feature grid ---------- */
.qa-grid {
  display: grid;
  grid-template-columns: repeat(2, 1fr);
  gap: 2.5rem;
  margin-top: 1.5rem;
  text-align: left;
}
.qa-card {
  background: transparent;
  border-top: 1px solid var(--line);
  border-radius: 0;
  padding: 1.75rem 0 0;
}
.qa-card .num {
  font-size: 1.6rem;
  font-style: italic;
  color: var(--orange-dark);
  margin-bottom: 0.5rem;
  display: block;
}
.qa-card p { margin-bottom: 0; color: var(--ink); font-size: 1.05rem; }

/* ---------- Lab values ---------- */
.values-illustration {
  display: block;
  max-width: 220px;
  width: 100%;
  height: auto;
  margin: 0 auto;
  transform: translateX(1mm);
}
.values-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: 2.5rem;
  margin-top: 1.5rem;
  text-align: left;
}
.value-card {
  background: transparent;
  border-top: 1px solid var(--line);
  border-radius: 0;
  padding: 1.75rem 0 0;
}
.value-card h3 { color: var(--orange-dark); font-size: 1.15rem; margin-bottom: 0.6rem; text-align: left; }
.value-card p { margin-bottom: 0; color: var(--ink); font-size: 1.02rem; text-align: left; }

/* ---------- Funding ---------- */
.funding-grid {
  display: flex;
  align-items: center;
  justify-content: center;
  flex-wrap: wrap;
  gap: 4rem;
  margin-top: 0.5rem;
}
.funder-logo { max-height: 160px; max-width: 420px; width: auto; height: auto; }
.funder-logo-simons { max-width: 240px; }
.funder-logo-hhmi { max-height: 185px; }

/* ---------- People ---------- */
.person-card {
  display: flex;
  gap: 3rem;
  align-items: flex-start;
  flex-wrap: wrap;
  margin-bottom: 4rem;
  text-align: left;
}
.person-card img.photo {
  width: 200px;
  height: 200px;
  object-fit: cover;
  border-radius: 8px;
  flex-shrink: 0;
}
.person-card .placeholder-photo {
  width: 200px;
  height: 200px;
  border-radius: 8px;
  background: var(--paper-alt);
  display: flex;
  align-items: center;
  justify-content: center;
  color: var(--ink-soft);
  font-size: 2.5rem;
  flex-shrink: 0;
}
.person-card .info { flex: 1; min-width: 280px; padding-top: 0.25rem; }
.person-card .role { color: var(--purple); font-size: 1rem; font-style: italic; margin-bottom: 0.6rem; }
.social-icon-link { display: inline-block; vertical-align: middle; margin-left: 0.15rem; opacity: 0.75; }
.social-icon-link:hover { opacity: 1; }
.social-icon { width: 18px; height: 18px; display: block; }

/* ---------- Publications ---------- */
.pub-list { list-style: none; margin-left: 0; text-align: left; }
.pub-list li {
  padding: 1.5rem 0;
  border-bottom: 1px solid var(--line);
  color: var(--ink);
}
.pub-list li:last-child { border-bottom: none; }
.pub-note { font-size: 0.9rem; color: var(--purple); font-style: italic; margin-top: 0.3rem; }
.pub-list a.read-link { font-size: 0.9rem; margin-left: 0.35rem; }

.scholar-link { margin-top: 1.75rem; text-align: center; }

.section-label {
  font-size: 0.95rem;
  font-style: italic;
  letter-spacing: 0.02em;
  color: var(--orange-dark);
  margin-bottom: 0.5rem;
}

.subtle-support {
  text-align: center;
  font-size: 0.8rem;
  color: var(--ink-soft);
  max-width: 560px;
  margin: 0 auto 2.5rem;
}
.subtle-support a { color: var(--ink-soft); text-decoration: underline; }
.subtle-support a:hover { color: var(--orange-dark); }

#join-us { scroll-margin-top: 7rem; }

/* ---------- Footer ---------- */
footer {
  padding: 3.5rem 0 3rem;
  color: var(--ink-soft);
  font-size: 0.9rem;
  background: transparent;
  border-top: 1px solid var(--line);
  text-align: center;
}
footer .wrap { display: flex; flex-direction: column; align-items: center; gap: 0.4rem; }
footer a { color: var(--ink-soft); }
footer a:hover { color: var(--orange-dark); }
.footer-logo { max-width: 320px; width: 100%; height: auto; margin-top: 1.25rem; }

/* ---------- Loading state for JS-rendered lists ---------- */
.loading-note { color: var(--ink-soft); font-style: italic; }

@media (max-width: 720px) {
  h1 { font-size: 2rem; }
  .hero { padding: 1.5rem 0 2rem; }
  .person-card { flex-direction: column; }
  .qa-grid { grid-template-columns: 1fr; }
  .pub-grid { grid-template-columns: 1fr; }
  .values-grid { grid-template-columns: 1fr; }
  .funding-grid { gap: 2rem; }
  .band { padding: 2.5rem 0; }
  section { margin-bottom: 3rem; }
  .nav-links { gap: 0.9rem; }
  .nav-links a { font-size: 1.05rem; }
  .dropdown-menu { right: 0; left: auto; min-width: 160px; }
  .research-figure img { width: 85vw; height: 85vw; max-width: 320px; max-height: 320px; }

  /* Shrink the home-page banner, logo, and nav so the header doesn't eat half the screen on phones */
  .banner .wrap { display: block; text-align: center; padding: 0.35rem 0.75rem; font-size: 0.72rem; }
  .banner .wrap a { margin-left: 0.35rem; white-space: nowrap; }
  .logo-row { padding-top: 0.05rem; }
  .logo-row img { height: 68px; }
  .header-tagline { font-size: 0.75rem; margin-top: -0.2rem; margin-bottom: -0.1rem; }
  nav.wrap { padding: 0.1rem 0.3rem 0.3rem; }
  .nav-links a { font-size: 0.95rem; }
  nav.wrap .nav-links { flex-wrap: nowrap; justify-content: center; gap: 0.45rem; }
  nav.wrap .nav-links a { font-size: 0.78rem; font-weight: 400; white-space: nowrap; }
  .hero h1 { font-size: 2.05rem; }
}